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Ukhaatherium is a now extinct species of mammal that lived during the upper Cretaceous about 84 to 72 million years ago in today's East Asia. It is known above all from the fossil locality Ukhaa Tolgod, Mongolia. An adult Ukhaatherium has an estimated weight of about 32g and bears several similarities to lipotyphlan insectivorans such as the tenrec.

Characteristics

Ukhaatherium nessovi, the type and only species for the genus, is known from multiple near-complete specimens. The most notable feature of the species is the presence of epipubic bones in the pelvic girdle, which have been lost in extant eutherian mammals. The loss of epipubic bones is associated with the evolution of prolonged gestation in eutherian mammalian reproduction. This means that Ukhaatherium may have had a short gestation period resulting in the birth of altricial young, like monotremes, marsupials, and extinct Mesozoic mammals such as multituberculates. This supports the hypothesis that the presence of epipubic bones is the primitive mammalian condition.

Despite some primitive skull and dental traits, the skeletons of Ukhaatherium and two other asioryctitheres, Asioryctes and Kennalestes, are classed as eutherians and show several similarities with the extant lipotyphlans. Derived eutherian characteristics include the restriction of the upper ankle joint to the parasagittal plane.

Fossil findings

Since the year 1990, joint expeditions of the American Museum of Natural History and the Mongolian Academy of Sciences have taken place in the Gobi desert of southern Mongolia in order to investigate upper Cretaceous and Tertiary vertebrates. More than 500 mammalian skulls (many of which with corresponding postcranial skeletons) were discovered at the Ukhaa Tolgod fossil field between 1993 and 1997, together with very well-preserved dinosaur skeletons, eggs, and embryos, birds, and lizards. Fossils of two species of basal eutherian mammal were found, including the first known Ukhaatherium specimens and a new Zalambdalestes specimen. These originate from the Djadochta Formation.

Paleobiology

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Systematics

Ukhaatherium is a genus from the family of the Asioryctitheria, which lived in the late Cretaceous. Ukhaatherium and Asioryctes form the subfamily Asioryctinae, while Kennalestes stands somewhat outside this group.

References

1. ^ Novacek, Michael J.; Rougier, Guillermo W.; Wible, John R.; McKenna, Malcolm C.; Dashzeveg, Demberelyin; Horovitz, Inés (1997-10-02). "Epipubic bones in eutherian mammals from the Late Cretaceous of Mongolia". Nature. 389 (6650): 483–486. doi:10.1038/39020. ISSN 1476-4687. PMID 9333234. S2CID 205026882.
2. ^ Horovitz, Inés (2003-12-24). "Postcranial skeleton of Ukhaatherium nessovi (Eutheria, Mammalia) from the Late Cretaceous of Mongolia". Journal of Vertebrate Paleontology. 23 (4): 857–868. doi:10.1671/2399-10. ISSN 0272-4634. S2CID 85809847.

v t e Eutheria
Kingdom: Animalia Phylum: Chordata Class: Mammalia Subclass: Theria Synapsida see Synapsida Cynodontia see Cynodontia Mammalia see Mammalia Eutheria see below↓ Eutheria
Eutheria †Acristatherium? †Ambolestes? †Bayshinoryctes †Bobolestes? †Cokotherium? †Durlstodon †Durlstotherium †Endotherium †Eomaia? †Hovurlestes †Hyotheridium? †Indoclemensia †Juramaia? †Microtherulum? †Montanalestes? †Murtoilestes? †Prokennalestes? †Sinodelphys? †Adapisoriculidae Adapisoriculus Afrodon Bustylus Deccanolestes? Garatherium Namibiodon Remiculus? Sahnitherium? Wyonycteris? †Asioryctitheria Asioryctes Bulaklestes Daulestes Kennalestes Sasayamamylos Uchkudukodon Ukhaatherium †Cimolestidae Aboletylestes Acmeodon Alveugena Ambilestes Avunculus Bagalestes Batodon? Betonnia Chacopterygus Cimolestes Didelphodus Gelastops Ilerdoryctes Maelestes? Naranius Paleotomus Pararyctes Procerberus Protentomodon Puercolestes Scollardius Tsaganius †Didymoconidae Archaeoryctes Ardynictis Didymoconus Hunanictis Kennatherium Zeuctherium †Horolodectidae Ferrequitherium Horolodectes †Leptictida Gypsonictopidae Gypsonictops Sikuomys Pseudorhyncocyonidae Diaphyodectes Leptictidium Phakodon Pseudorhyncocyon Leptictidae Amphigyion Blacktops Gallolestes Ictopidium Labes Lainodon Leptictis Leptonysson Megaleptictis Myrmecoboides Ongghonia Palaeictops Praolestes Prodiacodon Protictops Wania †Palaeoryctidae Aaptoryctes Aceroryctes Eoryctes Lainoryctes Nuryctes Ottoryctes Palaeoryctes Pinoryctes †Pantolesta Paroxyclaenidae Dulcidon Euhookeria Fratrodon Kiinkerishella Kopidodon Merialus Paraspaniella Paroxyclaenus Pugiodens Sororodon Spaniella Vulpavoides Pantolestidae Amaramnis Aphronorus Bessoecetor Bisonalveus Bogdia Bouffinomus Buxolestes Chadronia Coriphagus Cryptopithecus Dyspterna Galethylax Gobipithecus Kochictis Oboia Pagonomus Palaeosinopa Pantolestes Pentacodon Thelysia Todralestes †Taeniodonta Schowalteria Conoryctidae Conoryctella Conoryctes Huerfanodon Onychodectidae Onychodectes Stylinodontidae Ectoganus Psittacotherium Stylinodon Wortmania †Tillodontia Adapidium Azygonyx Basalina Benaius Chungchienia Dysnoetodon Esthonyx Franchaius Higotherium Interogale Kuanchuanius Lofochaius Megalesthonyx Meiostylodon Plesiesthonyx Plethorodon Simplodon Tillodon Trogosus Yuesthonyx †Zalambdalestidae Alymlestes Anchilestes Barunlestes Beleutinus Kulbeckia Prozalambdalestes Zalambdalestes Zhangolestes Zofialestes †Zhelestidae Alostera? Aspanlestes Avitotherium Azilestes? Borisodon Eoungulatum Eozhelestes Gallolestes Kharmerungulatum Khuduklestes? Lainodon Mistralestes? Oxlestes? Paranyctoides? Parazhelestes Sheikhdzheilia Sorlestes Valentinella? Zhalmousia Zhelestes †Pantodonta see Pantodonta Placentalia Afrotheria Boreoeutheria Xenarthra

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