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In anatomy, an apex (adjectival form: apical) is part of the shell of a mollusk. The apex is the pointed tip (the oldest part) of the shell of a gastropod, scaphopod, or cephalopod.

The apex is used in end-blown conches.

Gastropods

The word "apex" is most often used to mean the tip of the spire of the shell of a gastropod. The apex is the first-formed, and therefore the oldest, part of the shell. To be more precise, the apex would usually be where the tip of the embryonic shell or protoconch is situated, if that is still present in the adult shell (often it is lost or eroded away).

Coiled gastropod shells

The phrase apical whorls, or protoconch, means the whorls that constitute the embryonic shell at the apex of the shell, especially when this is clearly distinguishable from the later whorls of the shell, otherwise known as the teleoconch.

Comparison of the apical part and the whole shell of Otukaia kiheiziebisu:

Limpet-like gastropod shells

Where this feature is present, the space under the apex of a patellate or patelliform (limpet-like) gastropod shell is called the apical cavity.

Scaphopods

The apex of tusk shells is the small, open posterior end, and the opening itself is usually called the apical aperture.

Cephalopods

In orthocone cephalopods, the pointed end of the shell is called the apex, and shell growth is away from the apex and toward the aperture. The first chamber of the apex is sometimes called the protoconch.

Bivalves

The apex of a valve of a bivalve shell is the area around what is most commonly the umbo or beak. In some species the embryonic shell or prodissoconch may be present at the apex of each of the two valves.

References

1. Dance, S. Peter (2022). Shells. United Kingdom: Dorling Kindersley Limited. p. 250. ISBN 978-0-2415-1551-8.

v t e Cephalopod anatomy
Shell	Types External Ammonoid shell Argonautid eggcase Nautiloid shell Orthocone Internal Belemnoid guard Cirrate shell Cuttlebone Gladius (pen) Spirula shell Stylet Features External Aperture Apex Callus Lirae Periostracum Sculpture Spire Sutures Umbilicus Internal Body chamber Camerae Nacre Phragmocone Septa Siphuncle Whorls
Mantle & funnel	External anatomy Dermal structures Fins (wings) Flotation devices ("secondary fins") Funnel–mantle locking apparatus Tail Internal anatomy Branchial hearts Ctenidia (gills) Hepatopancreas (digestive gland) Ink sac and ink Needham's sac Nephridia ("kidneys") Nidamental glands Osphradium Pericardial glands
Head & limbs	Brachial crown Arms Hectocotylus Tentacles Carpus Dactylus Manus Suckers and hooks Buccal region Aptychus Beak Odontophore Radula Spadix and antispadix Occipital region Nuchal crest (occipital crest) Nuchal folds (occipital folds) Occipital membrane Olfactory organ Nuchal organ Other parts Eyes Statocysts
General	Chromatophores Photophores Nervous system Squid giant axon Squid giant synapse
Developmental stages: Spawn → Paralarva (Doratopsis stage) → Juvenile → Subadult → Adult • Egg fossils • Protoconch (embryonic shell)

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