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Actitis
Common sandpiper, Actitis hypoleucos
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Aves
Order:	Charadriiformes
Family:	Scolopacidae
Genus:	Actitis Illiger, 1811
Type species
Tringa hypoleucos (common sandpiper) Linnaeus, 1758
Species
Actitis hypoleucos Actitis macularius

Actitis is a small genus of waders, comprising just two very similar bird species.

Taxonomy

The genus Actitis was introduced in 1811 by the German zoologist Johann Illiger. The genus name is from Ancient Greek aktites, "coast-dweller" from akte, "coast". The type species is the common sandpiper. The genus is sister to the genus Tringa that contains the shanks and the tattlers.

The genus contains two species:

Genus Actitis – Illiger, 1811 – two species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Common sandpiper	Actitis hypoleucos (Linnaeus, 1758)	Eurasia	Size: Habitat: Diet:	LC
Spotted sandpiper	Actitis macularius (Linnaeus, 1766)	North America	Size: Habitat: Diet:	LC

Description

These are both small migratory waders, greyish brown on top and white underneath, with a distinctive stiff-winged flight low over the water. The plumages are very similar, apart from spotted sandpipers' distinctive breeding plumage, and suspected out-of-range vagrants must be carefully observed for identification to species.

Both species have short yellow or yellowish legs and a medium bill. These are not gregarious birds and are seldom seen in large flocks.

They nest on the ground, and their habitat is near fresh water. These birds forage on the ground or in water, picking up food by sight. They may also catch insects in flight. They eat insects, crustaceans and other invertebrates.

Actitis is part of the shank-tattler-phalarope clade and less closely related to the calidrid sandpipers. Based on the degree of DNA sequence divergence and putative shank and phalarope fossils from around the Oligocene/Miocene boundary some 23–22 million years ago, presumably Actitis diverged from its closest relatives in the Late Oligocene; given the much higher diversity of the prehistoric members of the group in Eurasia it is likely that they originated there, possibly being isolated as the remains of the Turgai Sea dried up, which happened just around this time.

The Late Pliocene fossil described as Actitis balcanica appears to be actually from some indeterminate charadriid.

Footnotes

1. Illiger, Johann Karl Wilhelm (1811). Prodromus systematis mammalium et avium (in Latin). Berolini : Sumptibus C. Salfeld. p. 262.
2. Jobling, James A (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. p. 31. ISBN 978-1-4081-2501-4.
3. Peters, James Lee, ed. (1934). Check-List of Birds of the World. Vol. 2. Cambridge, Massachusetts: Harvard University Press. p. 269.
4. Gibson, Rosemary; Baker, Allan (2012). "Multiple gene sequences resolve phylogenetic relationships in the shorebird suborder Scolopaci (Aves: Charadriiformes)". Molecular Phylogenetics and Evolution. 64 (1): 66–72. Bibcode:2012MolPE..64...66G. doi:10.1016/j.ympev.2012.03.008. PMID 22491071.
5. Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (July 2021). "Sandpipers, snipes, coursers". IOC World Bird List Version 11.2. International Ornithologists' Union. Retrieved 22 November 2021.
6. Paton et al. (2003), Thomas et al. (2004)
7. Mlíkovský (2002), Paton et al. (2003)
8. Boev, Z. 1998. Actitis balcanica sp. n. - a Late Pliocene Sandpiper (Aves: Scolopacidae) from Bulgaria. - Historia naturalis bulgarica, 9: 71–77.
9. Mlíkovský (2002)

References

• Mlíkovský, Jirí (2002): Cenozoic Birds of the World, Part 1: Europe. Ninox Press, Prague. ISBN 80-901105-3-8
• Paton, Tara A.; Baker, Allan J.; Groth, J. G.; Barrowclough, G. F. (2003). "RAG-1 sequences resolve phylogenetic relationships within charadriiform birds". Molecular Phylogenetics and Evolution. 29 (2): 268–278. Bibcode:2003MolPE..29..268P. doi:10.1016/S1055-7903(03)00098-8. PMID 13678682.
• Thomas, Gavin H.; Wills, Matthew A.; Székely, Tamás (2004). "A supertree approach to shorebird phylogeny". BMC Evolutionary Biology. 4: 28. doi:10.1186/1471-2148-4-28. PMC 515296. PMID 15329156. Supplementary Material

v t e Genera of shorebirds and their extinct allies
Kingdom: Animalia Phylum: Chordata Class: Aves Infraclass: Neognathae Clade: Neoaves Order: Charadriiformes
Charadriiformes incertae sedis †Charadriisimilis †Eupterornis? †Fluviatitavis? †Jiliniornis †"Morsoravis" †Nahmavis †Palintropus? †Sarjeantopodus †Scandiavis †Volgavis? †Laornithidae? Laornis †Graculavidae? Dakotornis Graculavus Palaeotringa Telmatornis? Charadrii See below ↓ Scolopaci See below ↓ Lari See below ↓
Charadrii incertae sedis †Cherevychnavis Burhinidae Burhinus Esacus †Genucrassum Pluvianellidae Pluvianellus Chionidae Chionis †Chionoides †Neilus Pluvianidae Pluvianus Pluvianidae Vanellinae †Dolicopterus? Erythrogonys Hoploxypterus Vanellus †Viator Charadriinae Anarhynchus Charadrius †Dorypaltus Elseyornis †Jilniornis Oreopholus Peltohyas Phegornis Pluvialis Thinornis Recurvirostridae Cladorhynchus Himantopus †Kashinia Recurvirostra Ibidorhynchidae Ibidorhyncha Haematopodidae Haematopus
Scolopaci incertae sedis †Hakawai †Scolopacimilis Jacanidae Actophilornis Hydrophasianus Irediparra Jacana †Janipes Metopidius Microparra †Nupharanassa Pedionomidae †Oligonomus Pedionomus Rostratulidae Nycticryphes Rostratula Scolopacidae Actitis Arenaria Bartramia Calidris Coenocorypha †Elorius Gallinago Limnocryptes Limnodromus Limosa †Mirolia Numenius †Nuntius †Paractitis †Parvelorius Phalaropus Prosobonia Scolopax Tringa †Villetus Xenus Thinocoridae Attagis Thinocorus
Lari Alcidae †Divisulcus †Hydrotherikornis †Miocepphus †Pseudocepphus Alcinae Alcini Alca Alle †Pinguinus Uria Synthliboramphini Synthliboramphus Cepphini Cepphus Brachyramphini Brachyramphus Fraterculinae Aethiini Aethia Ptychoramphus Fraterculini Cerorhinca Fratercula †Mancallinae Alcodes Mancalla Miomancalla Praemancalla Dromadidae Dromas Glareolidae †Becassius †Boutersemia †Mioglareola †Paractiornis †"Precursor"? Glareolinae Glareola Stiltia Glareolinae Cursorius Rhinoptilus Laridae Anous Chlidonias Chroicocephalus Creagrus †Feducciavis †Gaviota Gelochelidon Gygis Hydrocoloeus Hydroprogne Ichthyaetus †Laricola Larosterna Larus Leucophaeus †Limicolavis? †Limosavis Onychoprion Pagophila Phaetusa †Pseudosterna Rhodostethia Rynchops Rissa Sterna †Sternalara Sternula Thalasseus Xema Stercorariidae Stercorarius Turnicidae †Cerestenia †Eocliffia? Ortyxelos †Turnipax Turnix

• Actitis
• Bird genera
• Taxa named by Johann Karl Wilhelm Illiger